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28) with infaunal bivalves only forming 8% of the fauna (text-fig. 24). It ranges through 10 m of sediment in the mid-pectinatus Zone (textfig. 10) characterized by abundant fish scales and common Chondrites traces. Density is very high with an average of 199 specimens per sample. g. bed 49g, text-fig. 10). The abundant gastropods in this association may have eaten a bacterial mat or even benthic algae. Virtually the entire fauna lived on the sediment surface and the substrate was presumably very firm.

E9 Pseudorhytidopilus latissima: Protocardia morinica Association (6 samples, 1073 specimens, 16 species, 24% average fragmentation, H = 1-58) This is the most diverse of the dysaerobic associations with a fauna dominated by epifaunal forms (text-fig. 24), with the exception of the ubiquitous P. morinica (text-fig. 30). It is found in the mudstones and paper shales of the pectinatus and pallasioides Zones (text-figs 10 and 11). In the 42 WIGNALL: KIMMERIDGE CLAY BENTHIC PALAEOECOLOGY cm 0 % 20 VITrrrrrrri'iVi 1 1 1 1 T 1 f 11 1 1 1 1 1 1 1 iii Hi i111111• 1i111111111 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 E9 30.

Above bed 33 the change to more arenaceous sediments makes the numbering scheme inapplicable. TEXT-FIG. of the mudstone associations from Westbury, shallow infaunal filter feeding bivalves dominate the association (text-fig. 43). However, deep infaunal forms constitute 6-8%, whilst deposit feeders, mostly Nucinella birkelundi, make up 12-8% of the total. Association A14 occurs at the same level as associations Al and A4 at Wyke Regis, Dorset (textfig. 3) where organic-rich shales are better developed.

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